Is this a thing at places like Two Sigma, Citadel, and others? If it is, what does the process look like?
Maybe if you have reaearch experience or a phd. Otherwise almost impossible.
It happens at small funds
Depends on the group. Certain groups have smaller distinctions between the two roles.
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Usually not, no. If it does happen the process would be said applicant gaining relevant experience/credentials to be a quant and then going through the interview process just like anyone else.